Buffy Summer’s brother, Angel, is getting married to Drusilla, and while she is happy for him, she takes an instant dislike to her new brother-in-law, William Giles. The feeling is more than mutual. They walk out of each others lives without a second thought. When Dru and Angel die in a car accident, leaving custody of their six month old daughter to Buffy and William, they are forced to rethink their concept of "family" and their feelings for each other.